Osmotic pressure
For dilute solutions the van 't Hoff relation says osmotic pressure grows in proportion to dissolved particles. For ordinary salt water a handy simplification is about 0.7 bar for every 1,000 mg/L, so seawater near 35,000 mg/L has an osmotic pressure around 25 bar. The model uses that simplification throughout.
Assuming the membrane rejects nearly all salt, every bit of fresh water removed concentrates the rest. Brine salinity = feed salinity / (1 minus recovery). The net push at the outlet end is the feed pressure minus the brine's osmotic pressure, ignoring pressure losses along the vessel.
Why the brine matters
The concentrate leaving the plant still has to go somewhere: back to the sea through diffusers, into deep wells, or into evaporation ponds. High recovery saves feed water and pumping energy but makes the brine harder to handle and raises the risk of minerals scaling the membrane.